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Dr. Percy Alexander Hulley

Profile

Academic positionEmeritus
Research fieldsAnimal Systematics and Morphology,Animal Biochemistry and Physiology
Keywordsichthyology, lanternfishes, deep-sea

Current contact address

CountrySouth Africa
CityCape Town
InstitutionIziko - South African Museum
InstituteMarine Biology
Homepagewww.iziko.org.za/static/page/lantern-fish-myctophid-research

Host during sponsorship

Dr. Gerhard KrefftInstitut für Seefischerei, Bundesforschungsanstalt für Fischerei, Hamburg
Start of initial sponsorship01/09/1973

Program(s)

1973Humboldt Research Fellowship Programme

Publications (partial selection)

2023López-Pérez, C., Olivar, M.P., Tuset, V.M., Bernal, A. & Hulley, P.A.: Energy density of mesopelagic fishes from the Atlantic Ocean. In: Journal of Fish Biology, 102, 2023, 924-935
2023Mostarda, E., Harold, A.S., Sutton, T.T. & Hulley, P.A.: Identification Guide to the Mesopelagic Fishes of the western Indian Ocean. Food and Agriculture Organisation of the United Nations, 2023
2020Contreras, Tabit and Olivar, M Pilar and Gonz{\'a}lez-Gordillo, J Ignacio and Hulley, P Alexander: Feeding patterns of transforming and juvenile myctophids that migrate into neustonic layers. In: Marine Ecology Progress Series, 650, 2020, 239--252
2020Sutton, T.T. and Hulley, P.A. and Wienerroither, R. and Zaera-Perez, D. and Paxton, J.R.: Identification guide to the mesopelagic fishes of the central and south east {Atlantic} {Ocean}. Food and Agriculture Organisation of United Nations, 2020
2020L{\'o}pez-P{\'e}rez, Cristina and Olivar, M Pilar and Hulley, Percy A and Tuset, V{\'\i}ctor M: Length--weight relationships of mesopelagic fishes from the equatorial and tropical Atlantic waters: influence of environment and body shape. In: Journal of Fish Biology, 2020,
2019Contreras, Tabit and Olivar, M Pilar and Hulley, P Alexander and Fern{\'a}ndez de Puelles, M Luz: Feeding ecology of early life stages of mesopelagic fishes in the equatorial and tropical Atlantic. In: ICES Journal of Marine Science, 76, 2019, 673--689
2019Olivar, M Pilar and Bode, Antonio and L{\'o}pez-P{\'e}rez, Cristina and Hulley, P Alexander and Hern{\'a}ndez-Le{\'o}n, Santiago: Trophic position of lanternfishes (Pisces: Myctophidae) of the tropical and equatorial Atlantic estimated using stable isotopes. In: ICES Journal of Marine Science, 76, 2019, 649--661
2018Tuset, V{\'\i}ctor M and Olivar, M Pilar and Otero-Ferrer, Jose Lu{\'\i}s and L{\'o}pez-P{\'e}rez, C and Hulley, P Alexander and Lombarte, Antoni: Morpho-functional diversity in Diaphus spp.(Pisces: Myctophidae) from the central Atlantic Ocean: Ecological and evolutionary implications. In: Deep Sea Research Part I: Oceanographic Research Papers, 138, 2018, 46--59
2018Olivar, M Pilar and Contreras, Tabit and Hulley, P Alexander and Emelianov, Mikhail and L{\'o}pez-P{\'e}rez, Cristina and Tuset, V{\'\i}ctor and Castell{\'o}n, Arturo: Variation in the diel vertical distributions of larvae and transforming stages of oceanic fishes across the tropical and equatorial Atlantic. In: Progress in Oceanography, 160, 2018, 83--100
2017M.P. OLIVAR, P.A. HULLEY, A. CASTELLÓN, M. EMELIANOV, C. LÓPEZ, V.M. TUSET, T. CONTRERAS, B. MOLÍ Mesopelagic fishes across the tropical and equatorial Atlantic: Biogeographical and vertical patterns. In: Progress in Oceanography, 2017, 116-137
2017B.A. POLIDORO, G.M. RALPH, M. HARVEY, K.E. CARPENTER, ... P.A. HULLEY ... et al.: Red List of Marine Bony Fishes of the Eastern Central Atlantic. IUCN, 2017
2014G. DUHAMEL, P.A. HULLEY, R.CAUSSE, P. KOUBBI, M. VACCHI, P. PRUVOST, S. VIGETTA, J.-O. IRISSON, S. MORMÈDE, M. BELCHIER, A. DETTAI, H.W. DETRICH, J. GUTT, C.D. JONES, K.H. KOCK, L.J. LOPEZ ABELLAN, A. VAN DE PUTTE: Chapter 7. Biogeographic Patterns of Fish. In: C. DeBroyer, P. Koubbi, H.J. Griffiths, B. Raymond, C. d' Udekem d'Acoz, A. van de Putte, B. Danis, B. David, S. Grant, J. Gutt, C. Held, G. Hosie, F. Huettmann, A. Post, Y. Ropert-Coudert, Biogeographic Atlas of the Southern Ocean . Scientific Committee on Antarctic Research, 2014. 328-362
2014P. KOUBBI, R.J.M. CRAWFORD, N. ALLONCLE, N AMEZIANE, C. BARBRAUD, D. BESSON, C.A. BOST, K. DELORD, G. DUHAMEL, L. DOUGLASS, C. GUINET, G. HOSIE, P.A. HULLEY, J.-O. IRISSON, K.M. KOVACS, R. LESLIE, A. LOMBARD, A. MAKHADO, C. MARTINEZ, S. MORMÈDE, F. PENOT, P. PISTORIUS, P. PRUVOST, B. RAYMOND, E REUILLARD, J. RINGELSTEIN, T. SAMAAI, P. TIXIER, H.M. VERHEYE, S. VIGETTA, C. VON QUILLFELDT, H. WEIMERSKIRSCH Estimating the biodiversity of Planning Domain 5 (Marion and Prince Edward Islands – Del Cano – Crozet) for ecoregionalisation. WG-EMM-12/33. CCAMLR, 2014
2011P.A. HULLEY, G. DUHAMEL: Aspects of Lanternfish Distribution in the Kerguelen Plateau Region. In: G. Duhamel, D. Welsford, The Kerguelen Plateau: Marine Ecosystem and Fisheries. Société Française d' Ichtyologie, 2011. 183-195
2011P. KOUBBI, M. MOTEKI, G. DUHAMEL, A. GOURANT, P.A. HULLEY, R. O'DRISCOLL, T. ISHIMARU, P. PRUVOST, E. TAVERNIER, G. HOSIE: Ecoregionalization of myctophid fish in the Indian sector of the Southern Ocean: results from generalized dissimilarity models. In: Deep-Sea Research II,, 2011, 170-180
2011P. KOUBBI, P.A. HULLEY, P. PRUVOST, P. HENRI, J.-P. LABAT, V.A. WADLEY, H. DAISUKE, M. MOTEKI: Size distribution of meso- and bathypelagic fish in the Dumont d'Urville Sea (East Antarctica) during the CEAMARC surveys. In: Polar Science, 2011, 195-200
2011M. MOTEKI, P. KOUBBI, P. PRUVOST, E. TAVERNIER, P.A. HULLEY: Spatial distribution of pelagic fishes off Adélie and George V Land, East Antarctica in the austral summer 2008. In: Polar Science, 2011, 211-221
2010P. KOUBBI, C. OZOUF-COSTAZ, A. GOARANT, M. MOTEKI, P.A. HULLEY, R. CAUSSE, A. DETTAI, G. DUHAMEL, P. PRUVOST, E. TAVERNIER, A. POST, R.J. BEAMAN, S. RINTOUL, T. HIRAWAKE, T. HIRANO, T. ISHIMARU, M. RIDDLE, G.W. HOSIE: Estimating the biodiversity of the East Antarctic shelf and oceanic zone for ecoregionalisation: example of the ichthyofauna of the CEAMARC (Collaborative East Antarctic Marine Census) CAML surveys.. In: Polar Science, 2010, 115-133
2009P.A. HULLEY, G. DUHAMEL: A review of the lanternfish genus Bolinichthys Paxton, 1972 (Myctophidae). . In: Cybium, 2009, 259-304
2004C. PUSCH, P.A. HULLEY, K.H. KOCK: Community structure and feeding ecology of mesopelagic fishes in the slope waters of King George Island (South Shetland Islands, Antarctica). . In: Deep-sea Research I, 2004, 1685-1708
2002W.K. FLORENCE, P.A. HULLEY, B.A. STEWART, M.J. GIBBONS: Genetic and morphological variation of the lanternfish Lampanyctodes hectoris (Myctophiformes: Myctophidae) off southern Africa. In: South African Journal of Marine Science, 2002, 193-203
2002L.E. BECKLEY, P.A. HULLEY, P.H. SKELTON: Synoptic overview of marine ichthyology in South Africa. In: Marine and Freshwater Research, 2002, 99-105
2000J.R. PAXTON, P.A. HULLEY: Family Myctophidae (lanternfishes). In: A Checklist of the Fishes of the South China Sea (Randall, J.R. & Lim, K.K.P. eds.). In: The Raffles Bulletin of Zoology, 2000, 593-594
2000J.R. PAXTON, P.A. HULLEY: Family Neoscopelidae (blackchins). In: A Checklist of the Fishes of the South China Sea (Randall, J.R. & Lim, K.K.P. eds.). In: The Raffles Bulletin of Zoology, 2000, 592-593
2000M.E. ANDERSON, P.A. HULLEY: Functional ecosystems: the deep sea. . In: B.D. Durham, J.C.Pauw, Marine Biodiversity Status Report for South Africa at the end of the 20th Century. National Research Foundation, 2000. 20-25
1999?.?. ?????, ?.?. ??????????, ?.?. ????? ????? ??????? ?????? Astronesthes boulengeri (Astronestidae, Stomiiformes) ? ????????? ???? ????? ?????. In: Voprosÿ ikhtiologii, 1999, 581-594
1999J.R. PAXTON, P.A. HULLEY: Family Neoscopelidae. In: K.E. Carpenter, V.H. Niem, The Living Marine Resources of the Western Central Pacific. Volume 3. Batoid fishes, chimaeras and bony fishes part 1 (Elopidae to Linophrynidae). Food & Agriculture Organization of the United Nations, 1999. 1955-1956
1998K.H. KOCK, P.A. HULLEY: Fishes from Agassiz trawl catches. In: The expedition ANTARKTIS XIV/2 of RV 'Polarstern' in 1996/97 (Kattner, G. eds.). In: Berichte der Polarforschung, 1998, 23
1998V. SIEGEL, K.H. KOCK, P.A. HULLEY, C. PUSCH, L. DOËLLEFELD, M.G. WHITE: Investigation on the krill stock and the epipelagic zooplankton community in the Elephant Island region. In: Berichte der Polarforschung, 1998, 50-56
1998P.A. HULLEY, K.H. KOCK, C. PUSCH, M.G. WHITE: Mesopelagic fishes. In: The expedition ANTARKTIS XIV/2 of RV 'Polarstern' in 1996/97 (Kattner, G. eds.). In: Berichte der Polarforschung, 1998, 17-22
1998P.A. HULLEY: Preliminary Investigations on the Evolution of the Tribe Electronini (Myctophiformes, Myctophidae). In: G. di Prisco, E. Pisano, A.E. Clarke Fishes of Antarctica. A Biological Overview. Springer, 1998. 75-85
1998K.H. KOCK, L. DOËLLEFELD, P.A. HULLEY, H.-P. JÄHRIG, W. PETZEL, C. PUSCH, V. SIEGEL, M.G. WHITE: The composition of the fish fauna on the northern slope of King George Island. In: Berichte der Polarforschung, 1998, 14-17
1997P.A. HULLEY: Ecomorphology of Fishes (Luczkovich, J.J., Motta, P.J., Norton, S.F. & Liem, K.F. eds.). In: South African Journal of Zoology, 1997, 221-222
1997P.A. HULLEY, G. DUHAMEL: Lanternfishes (Myctophidae) collected during the 1971-PELAGIA cruise of R.V. 'Coriolis' in the south-west Pacific Ocean. In: Cybium, 1997, 299-317
1995P.A. HULLEY: Myctophidae. In: O. Okamura, Fishes collected by the R/V Shinkai Maru around Greenland. Japan Marine Fishery Resources Research Center, 1995. 100-105
1995P.A. HULLEY, J.R.E. LUTJEHARMS: The south-western limit for the warm-water, mesopelagic ichthyofauna of the Indo-West Pacific: lanternfishes (Myctophidae) as a case study. In: South African Journal of Marine Science, 1995, 185-205
1994M. STEHMANN, P.A. HULLEY: Gerhard Krefft: 30 March 1912 - 20 March 1993. In: Copeia, 1994, 558-564
1994P.A. HULLEY: Lanternfishes. In: J.R. Paxton, W.N. Eschmeyer, Encyclopedia of Fishes. University of New South Wales, 1994. 127-128
1993P.A. HULLEY: Go west, young man. In: 4th Indo-Pacific Fish Conference, Bangkok, Thailand (28 November - 4 December 1993), 1993, 99
1993G. DUHAMEL, P.A. HULLEY: Ichtyofaune méso et bathypelagique. In:. Campagnes SKALP 1987 et 1988 aux îles Kerguelen à bord des navires 'SKIF' et 'KALPER' (Duhamel, G. eds.). In: Publications de l'Institut Français pour la Recherche et la Technologie Polaires, 1993, 285-295
1992P.A.HULLEY: Upper-slope distributions of oceanic lanternfishes (family: Myctophidae). In: Marine Biology, 1992, 365-383
1990P.A. HULLEY: Mesopelagic ichthyofauna. In: Les rapports des campagnes à la mer à bord du 'Marion Dufresne' MD42/SIBEX du 3 janvier au 18 février 1985 (Mayzaud, P. eds.). In: Publications de la Mission de Récherche des Terres Australes et Antartiques Françaises, 1990, 173-181
1990P.A. HULLEY, G. DUHAMEL: Réport on prélevements RMT - fishes. In: Les rapports des campagnes à la mer aux îles Saint-Paul et Amsterdam . Arnaud, P. eds.). In: Publications de la Mission de Recherche des Terres Australes et Antarctiques Françaises, 1990, 80-89
1989P.A. HULLEY, P. CAMUS, G. DUHAMEL: Ichthyological results of Cruise MD-42/SIBEX-II. Part 2. Fishes from RMT-8 stations, with additional records of lanternfishes (Myctophidae, Osteichthyes) from the Indian Sector of the Southern Ocean. In: Cybium, 1989, 83-99
1989P.A. HULLEY: Lanternfishes (Osteichthyes, Myctophidae) from the region of Saint-Paul and Amsterdam islands. In: Mésogée, 1989, 40-58
1989P.A. HULLEY: Lanternfishes of the southern Benguela region. Part 2. Gymnoscopelus (Gymnoscopelus) bolini Andriashev in South African waters, with comments on the distribution of subantarctic myctophids in the eastern South Atlantic. In: Annals of the South African Museum, 1989, 221-240
1989P.A. HULLEY, J.R.E. LUTJEHARMS Lanternfishes of the southern Benguela region. Part 3. The pseudoceanic-oceanic interface. In: Annals of the South African Museum, 1989, 409-435
1989R.M. PROSCH, P.A. HULLEY, R.A. CRUICKSHANK: Mesopelagic fishes and some other forage organisms. In: A.I.L. Payne, R.J.M. Crawford, Oceans of life off southern Africa . Vlaeberg, 1989. 130-135
1988C.J.AUGUSTYN, P.A. HULLEY: In search of alternative deep-water resources. In: South African Shipping News and Fishing Industry Review, 1988, 57-59
1988P.A. HULLEY: Marvels of the deep. In: Musenews, 1988,
1987P.A. HULLEY, R.M. PROSCH Mesopelagic fish derivatives in the southern Benguela Upwelling Region. In: The Benguela and comparative ecosystems (Payne, A.I.L., Gulland, J.A. & Brink, K.H. eds.). In: South African Journal of Marine Science, 1987, 597-611
1986P.A. HULLEY: A taxonomic review of the lanternfish genus Triphoturus Fraser-Brunner, 1949 (Myctophidae, Osteichthyes). In: Annals of the South African Museum, 1986, 71-95
1986P.A. HULLEY: A.H.B. De Decker. In: South African Journal of Marine Science, 1986, 1-2
1986P.A. HULLEY: Family 123: Stylephoridae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 404
1986P.A. HULLEY: Family 131: Anoplogasteridae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 415
1986P.A. HULLEY: Family 25: Rajidae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 115-127
1986P.A. HULLEY: Family 26: Anacanthobatidae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 127-128
1986P.A. HULLEY: Family 70: Idiacanthidae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 234-235
1986P.A. HULLEY: Family 82: Anotopteridae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 278
1986P.A. HULLEY: Family 86: Myctophidae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes . Macmillan, 1986. 282-321
1986P.A. HULLEY: Family 87: Neoscopelidae. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 321-322
1986P.A. HULLEY: Family Myctophidae. In: W. Fischer, FAO Species Identification Sheets for Fishery Purposes: Fishing Areas 48, 58, 88 (Southern Ocean) . Food & Agriculture Organization of the United Nations, 1986. 316-322
1986P.A. HULLEY: Lanternfishes of the southern Benguela region. Part 1. Faunal complexity and distribution. In: Annals of the South African Museum, 1986, 227-249
1986P.A. HULLEY: Order Myctophiformes. In: M.M. Smith, P.C. Heemstra, Smiths' Sea Fishes. Macmillan, 1986. 282
1985P.A. HULLEY, G. KREFFT: A zoogeographic analysis of the fishes of the family Myctophidae (Osteichthyes, Myctophiformes) from the 1979-Sargasso Sea Expedition of R.V. Anton Dohrn. In: Annals of the South African Museum,, 1985, 19-53
1984P.A. HULLEY: Neoscopelidae. In: P.J.P. Whitehead, M.L.Bauchot, J.C. Hureau, J.N. Nielsen, E. Tortonese, Fishes of the North-eastern Atlantic and the Mediterranean. Volume 1. . UNESCO, 1984. 426-428
1984P.A. HULLEY: The South African Museum's Meiring Naude cruises. 14. Family Myctophidae (Osteichthyes, Myctophiformes). . In: Annals of the South African Museum, 1984, 53-96
1982P.A. HULLEY: Lightfish. In: Report on alternative resources in the Benguela System (de Villiers, G. eds.). In: Benguela Ecology Programme Report, 1982, 1-10
1981P.A. HULLEY: Results of the research cruises of FRV 'Walther Herwig' to South America. LVIII. Family Myctophidae (Osteichthyes, Myctophiformes). In: Archiv für Fischereiwissenschaft, 1981, 1-300
1980P.A. HULLEY: Family Myctophidae. In: Preliminary list of marine fishes for CLOFETA (Quéro, J.C. eds.). In: Marine Information (UNESCO), 1980, 22-23
1980P.A. HULLEY: Family Neoscopelidae. In: Preliminary list of marine fishes for CLOFETA (Quéro, J.C. eds.). In: Marine Information (UNESCO), 1980, 24
1977P.A. HULLEY, M. STEHMANN: The validity of Malacoraja Stehmann, 1970 (Chondrichthyes, Batoidei, Rajidae) and its phylogenetic significance. In: Annals of the South African Museum, 1977, 227-237
1973P.A. HULLEY: Interrelationships within the Anacanthobatidae (Chondrichthyes, Batoidei), with a description of the lectotype of Anacanthobatis marmoratus Von Bonde & Swart. In: Annals of the South African Museum, 1973, 131-158
1972P.A. HULLEY: A new species of southern African brevirajid skate (Chondrichthyes, Batoidei). In: Annals of the South African Museum, 1972, 253-263
1972P.A. HULLEY: A report on the mesopelagic fishes collected during the deep-sea cruises of R.S. 'Africana II', 1961-1966. In: Annals of the South African Museum, 1972, 197-236
1972P.A. HULLEY: Mesopelagic fishes from Vema Seamount (IK Station 52). In: Annals of the South African Museum, 1972, 237-244
1972P.A. HULLEY: Taxonomic Literature of Vertebrates: Blackwelder, R.E.. In: Bulletin of the Zoological Society of Southern Africa, 1972, 1-2
1972P.A. HULLEY: The family Gurgesiellidae (Chondrichthyes, Batoidei), with reference to Pseudoraja atlantica Bigelow & Schroeder. In: Copeia, 1972, 356-359
1972P.A. HULLEY: The origin, interrelationships and distribution of southern African Rajidae (Chondrichthyes, Batoidei). In: Annals of the South African Museum,, 1972, 1-103
1972P.A. HULLEY: The rare plectognath fish Macrorhamphosodes uradoi (Kamohara)(Triacanthodidae) in South African waters. In: Annals of the South African Museum, 1972, 191-195
1971P.A. HULLEY: Centrophorus squamosus (Bonneterre)(Chondrichthyes, Squalidae) in the eastern South Atlantic. In: Annals of the South African Museum, 1971, 265-270
1970P.A. HULLEY: An investigation of the Rajidae of the west and south coasts of southern Africa. In: Annals of the South African Museum, 1970, 151-220
1969P.A. HULLEY, R. RAU: A female Regalecus glesne from Cape Province, South Africa. In: Copeia, 1969, 835-839
1969P.A. HULLEY: A note on the spelling of Lacépède. In: Bulletin of the Zoological Society of Southern Africa, 1969, 1-2
1969P.A. HULLEY: The relationship between Raja miraletus Linnaeus and Raja ocellifera Regan based on a study of the clasper. In: Annals of the South African Museum, 1969, 137-147
1966P.A. HULLEY, M.J. PENRITH: Euprotomicroides zantedeschia, a new genus and species of pigmy dalatiid shark from South Africa. In: Bulletin of Marine Science, 1966, 222-229
1966P.A. HULLEY: The validity of Raja rhizacanthus Regan and Raja pullopunctata Smith based on a study of the clasper. In: . Annals of the South African Museum, 1966, 497-514
199N.V. PARIN, O.D. BORODULINA, P.A. HULLEY: A review of the Astronesthes boulengeri species group (Astronesthidae, Stomiiformes) with description of two new species. In: Journal of Ichthyology (USSR), 199, 557-570