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Profile
| Academic position | Full Professor |
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| Research fields | Evolution and Systematics of Plants and Fungi,Ecology and Biodiversity of Plants and Ecosystems |
| Keywords | pennate diatoms, Systematic, evolution, phylogeny, structure |
| Honours and awards | 2024: Award for special contribution to the development of scientific research activity at the University of St. Cyril and Methodius 2015: Award for significant contribution in environmental protection 2014: Most successful scientist on Ss Cyril and Methodius University for 2014 year. 2010: State award for the most important contribution to science |
Current contact address
| Country | Northern Macedonia |
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| City | Skopje |
| Institution | Ss. Cyril and Methodius University in Skopje |
| Institute | Institute of Biology, Department of Botany |
Host during sponsorship
| Dr. Richard M. Crawford | Alfred-Wegener-Institut für Polar- und Meeresforschung (AWI), Bremerhaven |
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| Start of initial sponsorship | 01/09/2007 |
Programme(s)
| 2007 | Humboldt Research Fellowship Programme |
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Publications (partial selection)
| 2026 | Levkov, Zlatko: A new Reimeria (Bacillariophyceae) species from lakes Ohrid and Prespa, Macedonia. In: Phytotaxa, 747, 2026, 190 – 198 |
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| 2026 | Rybak, Mateusz and Peszek, Łukasz and Górecka, Ewa and Zadrąg-Tęcza, Renata and Levkov, Zlatko: Morphological and molecular characterization of Halamphora lukasiewiczii sp. nov. (Bacillariophyceae, Amphipleuraceae), a new extremophilic diatom from petroleum seeps in the Polish Carpathians. In: PhytoKeys, 271, 2026, 219 – 239 |
| 2026 | Rybak, Mateusz and Levkov, Zlatko: The overlooked diversity within a well-known diatom: Unraveling the Luticola ventricosa complex. In: PhytoKeys, 274, 2026, 247 – 272 |
| 2025 | Saber, Abdullah A. and El-Sheekh, Mostafa M. and Salem, Olfat M. A. and Levkov, Zlatko and Cantonati, Marco and Alotaibi, Modhi O. and Saber, Hani: Adding to Our Knowledge on the Diatom and Green Algae Biodiversity of Egypt: Some New-to-Science, Poorly Known, and Newly Recorded Species. In: Water (Switzerland), 17, 2025, |
| 2025 | Al-Handal, Adil Y. and Levkov, Zlatko and Abdullah, Huda and Wulff, Angela: Species of the genus Gomphonema from Haqlan hot springs, Iraq, with the description of two new species. In: Diatom Research, 40, 2025, 93 – 119 |
| 2025 | Ljubešić, Zrinka and Jurina, Dunja and Pavlović, Tin and Živolić, Mario and Levkov, Zlatko and Mucko, Maja and Bosak, Sunčica and Gligora Udovič, Marija: The Croatian National Diatom Collection – an overview and future challenges. In: Acta Botanica Croatica, 84, 2025, 141 – 162 |
| 2025 | Heudre, David and Van de Vijver, Bart and Werum, Marcus and Lange-Bertalot, Horst and Moreau, Laura and Cantonati, Marco and Angeli, Nicola and Levkov, Zlatko and Wetzel, Carlos E.: Two new Amphora species from European rivers compared to morphologically similar species of the Amphora copulata complex (Catenulaceae, Bacillariophyceae). In: Botany Letters, 2025, |
| 2024 | Popović, Slađana and Vidaković, Danijela and Levkov, Zlatko and Jakovljević, Olga and Nikolić, Nataša and Subakov Simić, Gordana and Krizmanić, Jelena: THE DIVERSITY AND MORPHOLOGY OF THE GENUS LUTICOLA D.G.MANN (BACILLARIOPHYTA) FROM SUBAERIAL BIOFILMS OF CAVES IN SERBIA; [RAZNOLIKOST IN MORFOLOGIJA RODU KREMENASTIH ALG LUTICOLA D. G. MANN (BACILLARIOPHYTA) IZ SUBAERSKIH BIOFILMOV JAM V SRBIJI]. In: Acta Carsologica, 53, 2024, 83 – 98 |
| 2024 | Cid-Rodríguez, María and Cantonati, Marco and Angeli, Nicola and Bilous, Olena and Al-Harbi, Maha and Lange-Bertalot, Horst and Levkov, Zlatko and Piana, Lucia and Spitale, Daniel and Saber, Abdullah A.: The Diatom Genus Navicula in Spring Ecosystems with the Description of Navicula aquaesuavis sp. nov.. In: Water (Switzerland), 16, 2024, |
| 2023 | Vidaković, Danijela and Levkov, Zlatko and Krizmanić, Jelena and Beszteri, Bank and Gavrilović, Bojan and Ćirić, Miloš: A new small-celled naviculoid diatom species, Mayamaea pannonica sp. nov. (Bacillariophyceae) from soda pans in Serbia. In: Phycologia, 62, 2023, 268 – 276 |
| 2023 | Chudaev, Dmitry and Levkov, Zlatko: A new species of the genus Adlafia (Anomoeoneidaceae, Bacillariophyta) from Moscow Region (Russia) and North Macedonia. In: Phytotaxa, 629, 2023, 266 – 274 |
| 2023 | Solak, Cüneyt Nadir and Levkov, Zlatko and Hamilton, Paul B. and Wishkerman, Asher and Yilmaz, Elif and Bąk, Malgorzata and Peszek, Lukasz and Blanco, Saul: Description of Gomphonella saldanensis sp. nov. (Bacillariophyceae) from hydromagnesite stromatolites in Salda Lake, Turkey. In: Diatom Research, 38, 2023, 89 – 102 |
| 2023 | Kochoska, Hristina and Chardon, Cécile and Chonova, Teofana and Keck, François and Kermarrec, Lenaïg and Larras, Floriane and Lefrancois, Estelle and Rivera, Sinziana F. and Tapolczai, Kálmán and Vasselon, Valentin and Levkov, Zlatko and Rimet, Frédéric: Filling reference libraries with diatom environmental sequences: strengths and weaknesses. In: Diatom Research, 38, 2023, 103 – 127 |
| 2023 | Levkov, Zlatko and Zaova, Dušica: Halamphora ectorii sp. nov. – a new diatom species from saline spring Solenica in North Macedonia. In: Nova Hedwigia, 117, 2023, 255 – 263 |
| 2023 | Ács, Éva and Bíró, Tibor and Boros, Emil and Dobosy, Péter and Duleba, Mónika and Földi, Angéla and Kiss, Keve T. and Levkov, Zlatko and Orgoványi, Péter and Szén, Orsolya Pálné and Trábert, Zsuzsa and Vadkerti, Edit and Grigorszky, István: Halamphora taxa in Hungarian soda pans and shallow soda lakes detected via metabarcoding and microscopic analyses. In: Metabarcoding and Metagenomics, 7, 2023, 413 – 438 |
| 2023 | Sun, Geng and Levkov, Zlatko and Xiao, Weiyang and Moraj, Nikola and Udovič, Marija Gligora and Du, Jie and Chen, Qunlong and Plenković-Moraj, Anðelka: New diatom species Diploneis zhui sp. nov. from Jiuzhaigou National Nature Reserve, China. In: Phytotaxa, 629, 2023, 235 – 244 |
| 2023 | Udovič, Marija Gligora and Šušnjara, Mirela and Kulaš, Antonija and Goreta, Gordana and Arapov, Jasna and Levkov, Zlatko: One new species of Aneumastus D.G. Mann et Stickle (Bacillariophyceae) from Krka River, Croatia. In: Fottea, 23, 2023, 21 – 29 |
| 2023 | Al-Handal, Adil Y. and Levkov, Zlatko and Wulff, Angela: Rhoicosphenia johannessoniae (Bacillariophyceae), a new diatom from the Swedish coast of the Baltic Sea. In: Phytotaxa, 618, 2023, 69 – 78 |
| 2023 | Sun, Geng and Levkov, Zlatko and Xiao, Weiyang and Moraj, Nikola and UdOVič, Marija Gligora and Du, Jie and Chen, Qunlong and Plenković-Moraj, Andelka: Sellaphora jiuzhaienis sp. nov. (Bacillariophyceae)—A new diatom species from Jiuzhaigou National Nature Reserve, China. In: Phytotaxa, 626, 2023, 259 – 268 |
| 2023 | Wagner, Bernd and Tauber, Paul and Francke, Alexander and Leicher, Niklas and Binnie, Steven A. and Cvetkoska, Aleksandra and Jovanovska, Elena and Just, Janna and Lacey, Jack H. and Levkov, Zlatko and Lindhorst, Katja and Kouli, Katerina and Krastel, Sebastian and Panagiotopoulos, Konstantinos and Ulfers, Arne and Zaova, Dušica and Donders, Timme H. and Grazhdani, Andon and Koutsodendris, Andreas and Leng, Melanie J. and Sadori, Laura and Scheinert, Mirko and Vogel, Hendrik and Wonik, Thomas and Zanchetta, Giovanni and Wilke, Thomas: The geodynamic and limnological evolution of Balkan Lake Ohrid, possibly the oldest extant lake in Europe. In: Boreas, 52, 2023, 1 – 26 |
| 2023 | Levkov, Zlatko and Zaova, Dušica and Jüttner, Ingrid: Three new cymbelloid species in the genera Cymbella, Cymbopleura and Delicatophycus from the mineral spring Crvena Voda near Skopje, Republic of North Macedonia. In: Diatom Research, 38, 2023, 165 – 179 |
| 2022 | Vidaković, Danijela and Krizmanić, Jelena and Gavrilović, Bojan and Dojčinović, Biljana P. and Levkov, Zlatko and Dimkić, Ivica and Ćirić, Miloš: Artificial sandpit lake as a habitat of brackish diatom species. In: Botany Letters, 169, 2022, 360 – 369 |
| 2022 | Videska, Anastasija and Zaova, Dušica and Naumovska, Hristina and Kochoska, Hristina and Mitic-Kopanja, Danijela and Levkov, Zlatko: Cymbella biseriata sp. nov.-a unique living species of Cymbella Agardh (Bacillariophyceae) with biseriate striae. In: Phytotaxa, 561, 2022, 41 – 52 |
| 2022 | Udovič, Marija Gligora and Kulaš, Antonija and Šušnjara, Mirela and Arapov, Jasna and Blanco, Saul and Levkov, Zlatko: Cymbopleura amicula stat nov. et. nom. nov. (Bacillariophyceae)—a rare diatom species from a karst river in Croatia. In: Phytotaxa, 532, 2022, 139 – 151 |
| 2022 | Zaova, Dušica and Cvetkoska, Aleksandra and Wagner, Bernd and Francke, Alexander and Vogel, Hendrik and Levkov, Zlatko and Jovanovska, Elena: Diatom community responses to environmental change in Lake Ohrid (Balkan Peninsula) during the mid-Pleistocene Transition. In: Quaternary International, 622, 2022, 1 – 9 |
| 2022 | Kulaš, Antonija and Gligora Udovič, Marija and Tapolczai, Kálmán and Žutinić, Petar and Orlić, Sandi and Levkov, Zlatko: Diatom eDNA metabarcoding and morphological methods for bioassessment of karstic river. In: Science of the Total Environment, 829, 2022, |
| 2022 | Stanković, Igor and Gligora Udovič, Marija and Šušnjara, Mirela and Alegro, Antun and Levkov, Zlatko: Diversity and ecology of benthic diatoms in a mire with narrow pH gradient. In: Fottea, 22, 2022, 39 – 55 |
| 2022 | Jovanovska, Elena and Hauffe, Torsten and Stelbrink, Björn and Cvetkoska, Aleksandra and Levkov, Zlatko and Wagner, Bernd and Lacey, Jack H. and Ognjanova-Rumenova, Nadja and Hamilton, Paul B. and Brandenburg, Karen M. and Albrecht, Christian and Wilke, Thomas: Environmental filtering drives assembly of diatom communities over evolutionary time-scales. In: Global Ecology and Biogeography, 31, 2022, 954 – 967 |
| 2022 | Mašić, Ermin and Zaova, Dušica and Barudanović, Senka and Ognjanova-Rumenova, Nadja and Levkov, Zlatko: Rimocostatus bugojnicus gen.et. sp. nov. (Coscinodiscophyceae, Bacillariophyta)–a new fossil diatom genus from Gračanica, Bugojno palaeolake in Bosnia and Herzegovina. In: Diatom Research, 37, 2022, 255 – 262 |
| 2021 | Cvetkoska, Aleksandra and Jovanovska, Elena and Hauffe, Torsten and Donders, Timme H. and Levkov, Zlatko and Van de Waal, Dedmer B. and Reed, Jane M. and Francke, Alexander and Vogel, Hendrik and Wilke, Thomas and Wagner, Bernd and Wagner-Cremer, Friederike: Drivers of phytoplankton community structure change with ecosystem ontogeny during the Quaternary. In: Quaternary Science Reviews, 265, 2021, |
| 2021 | Schabetsberger, Robert and Jersabek, Christian D. and Levkov, Zlatko and Ehrenfellner, Bianca and Enoka, Laulu Fialelei and Faiilagi, Seumalo Afele: Limnological Characterization of Three Tropical Crater Lakes in the Archipelago of Samoa (Lanoto'o, Olomaga, Mataulano). In: Pacific Science, 75, 2021, 163 – 174 |
| 2021 | Kochoska, Hristina and Zaova, Dušica and Videska, Anastasija and Mitic-Kopanja, Danijela and Naumovska, Hristina and Wetzel, Carlos E. and Ector, Luc and Levkov, Zlatko: Sellaphora pelagonica (Bacillariophyceae), a new species from dystrophic ponds in the Republic of North Macedonia. In: Phytotaxa, 496, 2021, 121 – 133 |
| 2021 | Chudaev, Dmitry and Jüttner, Ingrid and Levkov, Zlatko: The genus Navicula (Bacillariophyceae, Naviculaceae) from the valley of the Adegoy River (Krasnodar Territory, Russia) and description of two new species. In: Phytotaxa, 494, 2021, 208 – 218 |
| 2020 | Wilke, Thomas and Hauffe, Torsten and Jovanovska, Elena and Cvetkoska, Aleksandra and Donders, Timme and Ekschmitt, Klemens and Francke, Alexander and Lacey, Jack H. and Levkov, Zlatko and Marshall, Charles R. and Neubauer, Thomas A. and Silvestro, Daniele and Stelbrink, Björn and Vogel, Hendrik and Albrecht, Christian and Holtvoeth, Jens and Krastel, Sebastian and Leicher, Niklas and Leng, Melanie J. and Lindhorst, Katja and Masi, Alessia and Ognjanova-Rumenova, Nadja and Panagiotopoulos, Konstantinos and Reed, Jane M. and Sadori, Laura and Tofilovska, Slavica and Van Bocxlaer, Bert and Wagner-Cremer, Friederike and Wesselingh, Frank P. and Wolters, Volkmar and Zanchetta, Giovanni and Zhang, Xiaosen and Wagner, Bernd: Deep drilling reveals massive shifts in evolutionary dynamics after formation of ancient ecosystem. In: Science Advances, 6, 2020, |
| 2020 | Mihalić, Katarina Caput and Galović, Ines and Wetzel, Carlos E. and Ector, Luc and Ilijanić, Nikolina and Miko, Slobodan and Wishkerman, Asher and Hamilton, Paul B. and Levkov, Zlatko: Envekadea vranaensis sp. Nov. a new diatom species (bacillariophyta) from the lacustrine holocene sediments of Lake Vrana, Croatia. In: Nova Hedwigia, 110, 2020, 1 – 9 |
| 2020 | Schabetsberger, Robert and Grapci-Kotori, Linda and Ibrahimi, Halil and Bilalli, Astrit and Levkov, Zlatko and Jersabek, Christian D. and Vorage, Marcel and Denoël, Mathieu and Kaiser, Roland and Pall, Karin and Eisendle, Ursula and Eder, Erich and Sadiku, Almedina: First limnological characterization of Lakes Leqinat and Drelaj in Bjeshkët e Nemuna National Park, Kosovo. In: Eco.mont, 13, 2020, 12 – 21 |
| 2020 | Moreno, David Fernández and Luís, Ana T. and Vidakovic, Danijela and Levkov, Zlatko and Castillo, Pedro M. Sánchez: Pinnularia baetica sp. nov. (Bacillarophyceae): Comparison with other panduriform species in the Mediterranean area. In: Phytotaxa, 435, 2020, 85 – 100 |
| 2020 | Zaova, Dušica and Cvetkoska, Aleksandra and Mitic-Kopanja, Danijela and Jovanovska, Elena and Ognjanova-Rumenova, Nadja and Levkov, Zlatko: Tertiarius minutulus sp. Nov. (Stephanodiscaceae, Bacillariophyta) – A new fossil diatom species from lake ohrid. In: European Journal of Taxonomy, 2020, 2020, 1 – 14 |
| 2019 | Kulichová, Jana and Neustupa, Jiří and Vrbová, Kateřina and Levkov, Zlatko and Kopalová, Kateřina: Asymmetry in Luticola species. In: Diatom Research, 34, 2019, 67 – 74 |
| 2019 | Zhang, Wei and Wang, Ting and Levkov, Zlatko and Jüttner, Ingrid and Ector, Luc and Zhou, Qi-Chao: Halamphora daochengensis sp. Nov., a new freshwater diatom species (bacillariophyceae) from a small mountain lake, Sichuan province, China. In: Phytotaxa, 404, 2019, 12 – 22 |
| 2019 | Wagner, Bernd and Vogel, Hendrik and Francke, Alexander and Friedrich, Tobias and Donders, Timme and Lacey, Jack H. and Leng, Melanie J. and Regattieri, Eleonora and Sadori, Laura and Wilke, Thomas and Zanchetta, Giovanni and Albrecht, Christian and Bertini, Adele and Combourieu-Nebout, Nathalie and Cvetkoska, Aleksandra and Giaccio, Biagio and Grazhdani, Andon and Hauffe, Torsten and Holtvoeth, Jens and Joannin, Sebastien and Jovanovska, Elena and Just, Janna and Kouli, Katerina and Kousis, Ilias and Koutsodendris, Andreas and Krastel, Sebastian and Lagos, Markus and Leicher, Niklas and Levkov, Zlatko and Lindhorst, Katja and Masi, Alessia and Melles, Martin and Mercuri, Anna M. and Nomade, Sebastien and Nowaczyk, Norbert and Panagiotopoulos, Konstantinos and Peyron, Odile and Reed, Jane M. and Sagnotti, Leonardo and Sinopoli, Gaia and Stelbrink, Björn and Sulpizio, Roberto and Timmermann, Axel and Tofilovska, Slavica and Torri, Paola and Wagner-Cremer, Friederike and Wonik, Thomas and Zhang, Xiaosen: Mediterranean winter rainfall in phase with African monsoons during the past 1.36 million years. In: Nature, 573, 2019, 256 – 260 |
| 2019 | Vidakovic, Danijela and Levkov, Zlatko and Hamilton, Paul B.: Neidiopsis borealis sp. nov., a new diatom species from the mountain shar planina, republic of North Macedonia. In: Phytotaxa, 402, 2019, 21 – 28 |
| 2019 | Cantonati, Marco and Angeli, Nicola and Lange-Bertalot, Horst and Levkov, Zlatko: New amphora and halamphora (bacillariophyta) species from springs in the northern apennines (Emilia-Romagna, Italy). In: Plant Ecology and Evolution, 152, 2019, 285 – 292 |
| 2019 | Levkov, Zlatko and Vidaković, Danijela and Cvetkoska, Aleksandra and Mitić-Kopanja, Danijela and Krstić, Svetislav and Van de Vijver, Bart and Hamilton, Paul B.: Observations of the genus muelleria (bacillariophyceae) from the Republic of north Macedonia. In: Plant Ecology and Evolution, 152, 2019, 293 – 312 |
| 2019 | Mihalic, Katarina Caput and Udovic, Marija Gligora and Galovic, Ines and Stankovic, Igor and Šušnjara, Mirela and Žutinic, Petar and Kulaš, Antonija and Špoljaric, Igor and Levkov, Zlatko: Tetramphora croatica sp. nov.-A new brackish-water species from lake vransko, Croatia. In: Phytotaxa, 401, 2019, 276 – 286 |
| 2018 | Stelbrink, Björn and Jovanovska, Elena and Levkov, Zlatko and Ognjanova-Rumenova, Nadja and Wilke, Thomas and Albrecht, Christian: Diatoms do radiate: evidence for a freshwater species flock. In: Journal of Evolutionary Biology, 31, 2018, 1969 – 1975 |
| 2018 | Cvetkoska, Aleksandra and Pavlov, Aleksandar and Jovanovska, Elena and Tofilovska, Slavica and Blanco, Saul and Ector, Luc and Wagner-Cremer, Friederike and Levkov, Zlatko: Spatial patterns of diatom diversity and community structure in ancient Lake Ohrid. In: Hydrobiologia, 819, 2018, 197 – 215 |
| 2017 | Kale, Aditi and Levkov, Zlatko and Karthick, Balasubramanian: Typification of two species of Luticola (Bacillariophyta) from aerophilic habitats of the Western Ghats, India. In: Phytotaxa, 298, 2017, 29 – 42 |
| 2016 | Zhang, X.S. and Reed, J.M. and Lacey, J.H. and Francke, A. and Leng, M.J. and Levkov, Z. and Wagner, B.: Complexity of diatom response to Lateglacial and Holocene climate and environmental change in ancient, deep and oligotrophic Lake Ohrid (Macedonia and Albania). In: Biogeosciences, 13, 2016, 1351 – 1365 |
| 2016 | Jovanovska, Elena and Cvetkoska, Aleksandra and Tofilovska, Slavica and Ognjanova-Rumenova, Nadja and Levkov, Zlatko: Description of a new fossil diatom genus, Cribrionella gen. nov. (Bacillariophyta) from quaternary sediments of Lake Ohrid. In: Phytotaxa, 252, 2016, 031 – 042 |
| 2016 | Jovanovska, Elena and Cvetkoska, Aleksandra and Hauffe, Torsten and Levkov, Zlatko and Wagner, Bernd and Sulpizio, Roberto and Francke, Alexander and Albrecht, Christian and Wilke, Thomas: Differential resilience of ancient sister lakes Ohrid and Prespa to environmental disturbances during the Late Pleistocene. In: Biogeosciences, 13, 2016, 1149 – 1161 |
| 2016 | Cvetkoska, Aleksandra and Jovanovska, Elena and Francke, Alexander and Tofilovska, Slavica and Vogel, Hendrik and Levkov, Zlatko and Donders, Timme H. and Wagner, Bernd and Wagner-Cremer, Friederike: Ecosystem regimes and responses in a coupled ancient lake system from MIS 5b to present: The diatom record of lakes Ohrid and Prespa. In: Biogeosciences, 13, 2016, 3147 – 3162 |
| 2016 | Pavlov, Aleksandar and Jovanovska, Elena and Wetzel, Carlos E. and Ector, Luc and Levkov, Zlatko: Freshwater Mastogloia (Bacillariophyceae) taxa from Macedonia, with a description of the epizoic M. sterijovskii sp. nov.. In: Diatom Research, 31, 2016, 85 – 112 |
| 2016 | Tofilovska, Slavica and Cvetkoska, Aleksandra and Jovanovska, Elena and Ognjanova-Rumenova, Nadja and Levkov, Zlatko: Two new fossil Cyclotella (Kützing) BrébBisson species from Lake Ohrid, Macedonia/Albania. In: Fottea, 16, 2016, 218 – 233 |
| 2015 | Cvetkoska, Aleksandra and Levkov, Zlatko and Reed, Jane M. and Wagner, Bernd and Panagiotopoulos, Konstantinos and Leng, Melanie J. and Lacey, Jack H.: Quaternary climate change and Heinrich events in the southern Balkans: Lake Prespa diatom palaeolimnology from the last interglacial to present. In: Journal of Paleolimnology, 53, 2015, 215 – 231 |
| 2015 | Cvetkoska, Aleksandra and Hamilton, Paul B. and Levkov, Zlatko: Surirella prespanensis sp. nov. and Surirella hinziae sp. nov., two new diatom (bacillariophyceae) species from ancient lake prespa (macedonia/albania/greece). In: Fottea, 15, 2015, 179 – 189 |
| 2015 | Jovanovska, Elena and Levkov, Zlatko and Edlund, Mark B.: The genus Diploneis ehrenberg ex cleve (Bacillariophyta) from lake Hövsgöl, Mongolia. In: Phytotaxa, 217, 2015, 201 – 248 |
| 2015 | Ognjanova-Rumenova, Nadja and Jovanovska, Elena and Cvetkoska, Aleksandra and Levkov, Zlatko: Two new Tertiarius (Bacillariophyta, Coscinodiscophyceae) species from Mariovo Neogene basin, Macedonia. In: Fottea, 15, 2015, 51 – 62 |
| 2014 | Jovanovska, Elena and Buczkó, Krisztina and Nakov, Teofil and Levkov, Zlatko: Diploneis transylvanica (Bacillariophyceae), a new diatom species from the Neogene fossil deposit in Romania. In: Nova Hedwigia, 99, 2014, 413 – 427 |
| 2014 | Zidarova, Ralitsa and Levkov, Zlatko and Van De Vijver, Bart: Four new Luticola taxa (Bacillariophyta) from Maritime Antarctica. In: Phytotaxa, 170, 2014, 155 – 168 |
| 2014 | Voigt, Ricarda and Levkov, Zlatko: Halamphora saxonica sp. nov. (Bacillariophyceae) from lakes in lignite-mining area in Saxony, Germany. In: Nova Hedwigia, 99, 2014, 213 – 222 |
| 2014 | Cvetkoska, Aleksandra and Levkov, Zlatko and Reed, Jane M. and Wagner, Bernd: Late Glacial to Holocene climate change and human impact in the Mediterranean: The last ca. 17ka diatom record of Lake Prespa (Macedonia/Albania/Greece). In: Palaeogeography, Palaeoclimatology, Palaeoecology, 406, 2014, 22 – 32 |
| 2014 | Zhang, Xiaosen and Reed, Jane and Wagner, Bernd and Francke, Alexander and Levkov, Zlatko: Lateglacial and Holocene climate and environmental change in the northeastern Mediterranean region: Diatom evidence from Lake Dojran (Republic of Macedonia/Greece). In: Quaternary Science Reviews, 103, 2014, 51 – 66 |
| 2014 | Peng, Yumei and Rioual, Patrick and Levkov, Zlatko and Wiliams, David M and Jin, Zhangdong: Morphology and ultrastructure of Hippodonta qinghainensis sp. Nov. (bacillariophyceae), a new diatom from lake Qinghai, China. In: Phytotaxa, 186, 2014, 061 – 074 |
| 2014 | Tofilovska, Slavica and Wetzel, Carlos E. and Ector, Luc and Levkov, Zlatko: Observation on Achnanthes Bory sensu stricto (Bacillariophyceae) from subaerial habitats in Macedonia and comparison with the type material of A. coarctata (Brébisson ex W. Smith) Grunow, A. coarctata var. sinaensis Hustedt and A. intermedia Kützing. In: Fottea, 14, 2014, 15 – 42 |
| 2014 | Cvetkoska, Aleksandra and Hamilton, Paul B. and Ognjanova-Rumenova, Nadja and Levkov, Zlatko: Observations of the genus Cyclotella (Kützing) Brébisson in ancient lakes Ohrid and Prespa and a description of two new species c. paraocellata sp. nov. and C. prespanensis sp. nov. In: Nova Hedwigia, 98, 2014, 313 – 340 |
| 2014 | Cvetkoska, Aleksandra and Levkov, Zlatko and Hamilton, Paul B.: Surirella subrotunda sp. nov. and Surirella parahelvetica sp. nov., two new diatom (Bacillariophyta) species from Lake Prespa, Macedonia. In: Phytotaxa, 156, 2014, 145 – 155 |
| 2014 | Wagner, B. and Wilke, T. and Krastel, S. and Zanchetta, G. and Sulpizio, R. and Reicherter, K. and Leng, M.J. and Grazhdani, A. and Trajanovski, S. and Francke, A. and Lindhorst, K. and Levkov, Z. and Cvetkoska, A. and Reed, J.M. and Zhang, X. and Lacey, J.H. and Wonik, T. and Baumgarten, H. and Vogel, H.: The SCOPSCO drilling project recovers more than 1.2 million years of history from Lake Ohrid. In: Scientific Drilling, 2014, 19 – 29 |
| 2014 | Cvetkoska, Aleksandra and Levkov, Zlatko and Hamilton, Paul B. and Potapova, Marina: The biogeographic distribution of Cavinula (Bacillariophyceae) in North America with the descriptions of two new species. In: Phytotaxa, 184, 2014, 181 – 207 |
| 2014 | Mitić-Kopanja, Danijela and Wetzel, Carlos E. and Ector, Luc and Levkov, Zlatko: Two new Gomphonema EHRENBERG (Bacillariophyceae) species from Macedonia and comparison with type material of G. brebissonii KÜTZING. In: Fottea, 14, 2014, 149 – 160 |
| 2013 | Jovanovska, Elena and Buczkó, Krisztina and Ognjanova-Rumenova, Nadja G. and Nakov, Teofil and Levkov, Zlatko: Identity and typification of Diploneis ostracodarum, Diploneis budayana and Diploneis praeclara (Bacillariophyta). In: Phytotaxa, 137, 2013, 15 – 26 |
| 2013 | Jovanovska, Elena and Nakov, Teofil and Levkov, Zlatko: Observations of the genus Diploneis from Lake Ohrid, Macedonia. In: Diatom Research, 28, 2013, 237 – 262 |
| 2013 | Pavlov, Aleksandar and Levkov, Zlatko and Williams, David M. and Edlund, Mark B.: Observations on Hippodonta (Bacillariophyceae) in selected ancient lakes. In: Phytotaxa, 90, 2013, 1 – 53 |
| 2012 | Levkov, Zlatko and Williams, David M.: Checklist of diatoms (Bacillariophyta) from Lake Ohrid and Lake Prespa (Macedonia), and their watersheds. In: Phytotaxa, 45, 2012, 1 – 76 |
| 2012 | Zlatko Levkov, David M. Wiliams: Checklist of diatoms (Bacillariophyta) from Lake Ohrid and Lake Prespa (Macedonia), and their watersheds. . In: Phytotaxa, 2012, 1-76 |
| 2012 | Zlatko Levkov, Cvetkoska Aleksandra, Jane M. Reed: Lake Ohrid diatoms as palaeoclimate indicators of climate change during the last glacial-interglacial cycle. In: Diatom Monographs, 2012, 1-216 |
| 2011 | Levkov, Zlatko and Williams, David M.: Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia. In: Phytotaxa, 30, 2011, 1 – 41 |
| 2011 | Zlatko Levkov and David M. WIlliams: Fifteen new diatom (Bacillariophyta) species from Lake Ohrid, Macedonia. . In: Phytotaxa, 2011, 1-41 |
| 2011 | Ács, Éva and Ector, Luc and Kiss, Keve T. and Cserháti, Csaba and Morales, Eduardo A. and Levkov, Zlatko: Morphological observations and emended description of amphora micrometra from the bolivian altiplano, South America. In: Diatom Research, 26, 2011, 199 – 212 |
| 2011 | Eva Ács, Luc Ector, Keve T. Kiss, C. Cserháti, Eduardo A. Morales and Zlatko Levkov Morphological observations on Amphora micrometra Giffen from the Bolivian Altiplano, South America with emended diagnosis. . In: Diatom Research , 2011, 199-212 |
| 2010 | Zlatko Levkov and Luc Ector: A comparative study of Reimeria species (Bacillariophyceae). In: Nova Hedwigia , 2010, 469-489 |
| 2010 | Levkov, Zlatko and Ector, Luc: A comparative study of Reimeria species (Bacillariophyceae). In: Nova Hedwigia, 90, 2010, 469 – 489 |
| 2010 | Levkov, Zlatko and Caput Mihalić, Katarina and Ector, Luc: A taxonomical study of Rhoicosphenia GRUNOW (Bacillariophyceae) with a key for identification of selected taxa. In: Fottea, 10, 2010, 145 – 200 |
| 2010 | Zlatko Levkov, Katarina Caput-Mihalic and Luc Ector: A taxonomical study of Rhoicosphenia Grunow (Bacillariophyceae) with a key for identification of taxa. . In: Fottea , 2010, 1-56 |
| 2010 | Reed, J.M. and Cvetkoska, A. and Levkov, Z. and Vogel, H. and Wagner, B.: The last glacial-interglacial cycle in Lake Ohrid (Macedonia/Albania): Testing diatom response to climate. In: Biogeosciences, 7, 2010, 3083 – 3094 |
| 2010 | Jane M. Reed, Cvetkoska Aleksandra, Zlatko Levkov, H. Vogel, &. Bernd Wagner. The last glacial-interglacial cycle in Lake Ohrid (Macedonia/Albania): testing diatom response to climate. . In: Biogeosciences , 2010, 3083-3094 |
| 2009 | Zlatko Levkov: Amphora sensu lato. . In: H. Lange-Bertalot (ed.), Diatoms of Europe, Diatoms of the European Inland waters and comparable habitats. Vol. 5. A.R.G. Gantner Verlag K.G., 2009 |
| 2009 | Mark B. Edlund, Avery L.C. Shinneman, & Zlatko Levkov: Diatom biodiversity in Mongolia: A new amphoroid diatom from saline lakes in western Mongolia, Amphora soninkhishigae sp. nov. . In: Acta Botanica Croatica , 2009, 251-262 |
| 2009 | Pavlov, Aleksandar and Nakov, Teofil and Levkov, Zlatko and Furey, Paula and Lowe, Rex and Ector, Luc: Luticola grupcei (Bacillariophyceae) - A new freshwater diatom from Mountain Baba (Macedonia) and Great Smoky Mountains National Park (U.S.A.): comparison with the type material of L. goeppertiana (Bleisch) D.G.Mann. In: Nova Hedwigia, 89, 2009, 147 – 164 |
| 2009 | Aleksandar Pavlov, Teofil Nakov, Zlatko Levkov., Rex Lowe., Paula Furey & Luc Ector: Luticola grupcei (Bacillariophyceae) - a new freshwater diatom from Mountain Baba (Macedonia) and Great Smoky Mountains National Park (USA): comparison with the type material of L. goeppertiana (Bleisch) D.G. Mann. . In: Nova Hedwigia, 2009, 147-164 |
| 2009 | Nikolaos P. Nikolaidis, Aristomenis P. Karageorgis, Vasilios Kapsimalis, Paraskevi Drakopoulou, Nikolaos Skoulikidis, Horst Behrendt, Zlatko Levkov: Management of nutrient emissions of Axios River catchment: Their effect in the coastal zone of Thermaikos Gulf, Greece. In: Ecological Modelling, 2009, 383-396 |
| 2009 | Nikolaidis, Nikolaos P. and Karageorgis, Aristomenis P. and Kapsimalis, Vasilios and Drakopoulou, Paraskevi and Skoulikidis, Nikolaos and Behrendt, Horst and Levkov, Zlatko: Management of nutrient emissions of Axios River catchment: Their effect in the coastal zone of Thermaikos Gulf, Greece. In: Ecological Modelling, 220, 2009, 383 – 396 |
| 2008 | Levkov, Zlatko and Edlund, Mark B. and Nakov, Teofil: Identity and typification of Navicula hasta (Bacillariophyceae). In: Phycological Research, 56, 2008, 46 – 57 |
| 2008 | Zlatko Levkov, Teofil Nakov: Rhoicosphenia tenuis a new diatom species from Lake Ohrid. In: Diatom Research , 2008, 377-388 |
| 2008 | Levkov, Zlatko and Nakov, Teofil: Rhoicosphenia tenuis, a new diatom species from lake ohrid. In: Diatom Research, 23, 2008, 377 – 388 |
| 2008 | Zlatko Levkov, Mark B. Edlund and Teofil Nakov : The identity and typification of Navicula hasta (Bacillariophyceae). . In: Phycological Research , 2008, 46-57 |
| 2007 | Krstić, Svetislav and Svirčev, Zorica and Levkov, Zlatko and Nakov, Teofil: Selecting appropriate bioindicators regarding Water Framework Directive guidelines for freshwaters - A Macedonian experience. In: International Journal on Algae, 9, 2007, 41 – 63 |
| 2006 | Edlund, Mark B. and Brant, Lynn A. and Levkov, Zlatko and Nakov, Teofil: An emended description of decussata (patrick) lange-bertalot & metzeltin that includes protoplast organization and detailed valve and cingulum ultrastructure. In: Diatom Research, 21, 2006, 269 – 280 |
| 2006 | Levkov, Zlatko and Nakov, Teofil and Metzeltin, Ditmar: New species and combination from the genus sellaphora mereschkowsky from Macedonia. In: Diatom Research, 21, 2006, 297 – 312 |
| 2006 | Levkov, Zlatko and Williams, David M.: Two new species of diatoma from lakes ohrid and prespa, Macedonia. In: Diatom Research, 21, 2006, 281 – 296 |
| 2005 | Levkov, Zlatko and Krstic, Svetislav and Nakov, Teofil and Melovski, Ljupco: Diatom assemblages on Shara and Nidze Mountains, Macedonia. In: Nova Hedwigia, 81, 2005, 501 – 537 |
| 2002 | Krstic, Svetislav and Duma, Aleksej and Janevska, Biljana and Levkov, Zlatko and Nikolova, Ksenija and Noveska, Marina: Diatoms in forensic expertise of drowning - A Macedonian experience. In: Forensic Science International, 127, 2002, 198 – 203 |
| 2002 | Levkov, Z. and Krstic, S.: Use of algae for monitoring of heavy metals in the River Vardar, Macedonia. In: Mediterranean Marine Science, 3, 2002, 99 – 112 |